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The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
family,
Cervidae Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
, and one of the largest terrestrial
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s in its native range of North America and
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and
East Asia East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea and ...
. The common name of "elk" is open to confusion, as "elk" is the name used in
British English British English (BrE, en-GB, or BE) is, according to Lexico, Oxford Dictionaries, "English language, English as used in Great Britain, as distinct from that used elsewhere". More narrowly, it can refer specifically to the English language in ...
for the larger ''
Alces alces The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult ma ...
'', with similar names used by other European languages (German ''Elch'', Swedish ''älg'', and French ''élan''). In North America, the common name for ''Alces alces'' is "moose". The name "wapiti" is sometimes used for ''Cervus canadensis'', which derives from the
Shawnee The Shawnee are an Algonquian-speaking indigenous people of the Northeastern Woodlands. In the 17th century they lived in Pennsylvania, and in the 18th century they were in Pennsylvania, Ohio, Indiana and Illinois, with some bands in Kentucky a ...
and
Cree The Cree ( cr, néhinaw, script=Latn, , etc.; french: link=no, Cri) are a Indigenous peoples of the Americas, North American Indigenous people. They live primarily in Canada, where they form one of the country's largest First Nations in Canada ...
word ''waapiti'', meaning 'white rump'. Elk range in forest and forest-edge habitat, feeding on grasses, plants, leaves, and bark. Male elk have large
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...
s which they
shed A shed is typically a simple, single-story roofed structure that is used for hobbies, or as a workshop in a back garden or on an allotment. Sheds vary considerably in their size and complexity of construction, from simple open-sided ones de ...
each year. Males also engage in ritualized mating behaviors during the rut, including posturing, antler wrestling (sparring), and ''bugling'', a loud series of vocalizations that establishes dominance over other males and attracts females. Although it is currently native to North America and central/eastern Asia, it had a much wider distribution in the past. Populations were present across Eurasia into Western Europe during the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division of ...
and survived into the early
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
in southern Sweden and the Alps; the extinct
Merriam's elk The Merriam's elk (''Cervus canadensis merriami'') is an extinct subspecies of elk once found in the arid lands of the southwestern United States (in Arizona, New Mexico and Texas), as well as in Mexico. Uncontrolled hunting and cattle grazing si ...
subspecies ranged into Mexico. The elk has
adapted In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
well to countries where it has been introduced, including
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
. Its adaptability may in fact threaten
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
species and the ecosystems into which it has been introduced. Elk are susceptible to a number of
infectious disease An infection is the invasion of tissues by pathogens, their multiplication, and the reaction of host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmissible disease or communicable dise ...
s, some of which can be transmitted to
livestock Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to animals ...
. Efforts to eliminate infectious diseases from elk populations, largely by
vaccination Vaccination is the administration of a vaccine to help the immune system develop immunity from a disease. Vaccines contain a microorganism or virus in a weakened, live or killed state, or proteins or toxins from the organism. In stimulating ...
, have had mixed success. Some cultures revere the elk as having spiritual significance. In parts of Asia, antlers and their
velvet Weave details visible on a purple-colored velvet fabric Velvet is a type of woven tufted fabric in which the cut threads are evenly distributed, with a short pile, giving it a distinctive soft feel. By extension, the word ''velvety'' means ...
are used in
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
s. Elk are hunted as a game species. Their meat is leaner and higher in
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
than
beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantity ...
or
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
. Elk were long believed to belong to a
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of the European
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
(''Cervus elaphus''), but evidence from many
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
genetic studies beginning in 1998 shows that the two are distinct
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. Key morphological differences that distinguish ''C. canadensis'' from ''C. elaphus'' are the former's wider rump patch and paler-hued antlers.


Naming and etymology

By the 17th century, ''Alces alces'' (called "elk" in Europe) had long been
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
from the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
, and the meaning of the word "elk" to English-speakers became rather vague, acquiring a meaning similar to "large deer". The name ''wapiti'' is from the
Shawnee The Shawnee are an Algonquian-speaking indigenous people of the Northeastern Woodlands. In the 17th century they lived in Pennsylvania, and in the 18th century they were in Pennsylvania, Ohio, Indiana and Illinois, with some bands in Kentucky a ...
and
Cree The Cree ( cr, néhinaw, script=Latn, , etc.; french: link=no, Cri) are a Indigenous peoples of the Americas, North American Indigenous people. They live primarily in Canada, where they form one of the country's largest First Nations in Canada ...
word ''waapiti'' (in
Cree syllabics Cree syllabics are the versions of Canadian Aboriginal syllabics used to write Cree dialects, including the original syllabics system created for Cree and Ojibwe. There are two main varieties of syllabics for Cree: Western Cree syllabics and E ...
: or ), meaning "white rump". There is a subspecies of wapiti in
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
called the Altai wapiti (''Cervus canadensis sibiricus''), also known as the Altai maral. According to the ''
Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the first and foundational historical dictionary of the English language, published by Oxford University Press (OUP). It traces the historical development of the English language, providing a com ...
'', the etymology of the word "elk" is "of obscure history". In
Classical Antiquity Classical antiquity (also the classical era, classical period or classical age) is the period of cultural history between the 8th century BC and the 5th century AD centred on the Mediterranean Sea, comprising the interlocking civilizations of ...
, the European ''Alces alces'' was known as and , words probably borrowed from a
Germanic language The Germanic languages are a branch of the Indo-European language family spoken natively by a population of about 515 million people mainly in Europe, North America, Oceania and Southern Africa. The most widely spoken Germanic language, Engli ...
or another language of northern Europe. By the 8th century, during the
Early Middle Ages The Early Middle Ages (or early medieval period), sometimes controversially referred to as the Dark Ages, is typically regarded by historians as lasting from the late 5th or early 6th century to the 10th century. They marked the start of the Mi ...
, the moose was known as derived from the
Proto-Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the reconstructed proto-language of the Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic branc ...
: ''*elho-'', ''*elhon-'' and possibly connected with the . Later, the species became known in
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English p ...
as ''elk'', ''elcke'', or ''elke'', appearing in the Latinized form ''alke'', with the spelling ''alce'' borrowed directly from . Noting that ''elk'' "is not the normal phonetic representative" of the Old English ''elch'', the ''Oxford English Dictionary'' derives ''elk'' from , itself from . The American ''Cervus canadensis'' was recognized as a relative of the red deer (''Cervus elaphus'') of Europe, and so ''Cervus canadensis'' was referred to as "red deer"
Richard Hakluyt Richard Hakluyt (; 1553 – 23 November 1616) was an English writer. He is known for promoting the English colonization of North America through his works, notably ''Divers Voyages Touching the Discoverie of America'' (1582) and ''The Pri ...
, in his 1584 ''
Discourse Concerning Western Planting ''Discourse Concerning Western Planting'' was a document written by Richard Hakluyt in 1584. This document was written to convince Queen Elizabeth I to support the colonization schemes of Walter Raleigh and to encourage English merchants and ge ...
'', mentioned the continent's plentiful red deer (). Similarly, John Smith's 1616 '' A Description of New England'' referred to red deer. Sir William Talbot's 1672 English translation of
John Lederer John Lederer was a 17th-century German physician and an explorer of the Appalachian Mountains. He and the members of his party became the first Europeans to crest the Blue Ridge Mountains (1669) and the first to see the Shenandoah Valley and the ...
's Latin ''Discoveries'' likewise called the species "Red Deer", but noted in
parentheses A bracket is either of two tall fore- or back-facing punctuation marks commonly used to isolate a segment of text or data from its surroundings. Typically deployed in symmetric pairs, an individual bracket may be identified as a 'left' or 'r ...
that they were "for their unusual largeness improperly termed Elks by ignorant people". Both
Thomas Jefferson Thomas Jefferson (April 13, 1743 – July 4, 1826) was an American statesman, diplomat, lawyer, architect, philosopher, and Founding Fathers of the United States, Founding Father who served as the third president of the United States from 18 ...
's 1785 ''
Notes on the State of Virginia ''Notes on the State of Virginia'' (1785) is a book written by the American statesman, philosopher, and planter Thomas Jefferson. He completed the first version in 1781 and updated and enlarged the book in 1782 and 1783. It originated in Jeffers ...
'' and
David Bailie Warden David Bailie Warden was a republican insurgent in the Irish Rebellion of 1798 and, in later exile, a United States consul in Paris. While in American service Watson protested the corruption of diplomatic service by the "avaricious" spirit of com ...
's 1816 ''Statistical, Political, and Historical Account of the United States'' used "red deer" to refer to ''Cervus canadensis''..


Taxonomy

Members of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Cervus ''Cervus'' is a genus of deer that primarily are native to Eurasia, although one species occurs in northern Africa and another in North America. In addition to the species presently placed in this genus, it has included a whole range of other sp ...
'' (and hence early relatives or possible ancestors of the elk) first appear in the
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
record 25 million years ago, during the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
in
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
, but do not appear in the North American fossil record until the early
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
. The extinct
Irish elk The Irish elk (''Megaloceros giganteus''), also called the giant deer or Irish deer, is an extinct species of deer in the genus ''Megaloceros'' and is one of the largest deer that ever lived. Its range extended across Eurasia during the Pleisto ...
(''Megaloceros'') was not a member of the genus ''Cervus'' but rather the largest member of the wider deer family (Cervidae) known from the fossil record. Until recently, red deer and elk were considered to be one species, ''Cervus elaphus'', with over a dozen subspecies. But
mitochondria A mitochondrion (; ) is an organelle found in the Cell (biology), cells of most Eukaryotes, such as animals, plants and Fungus, fungi. Mitochondria have a double lipid bilayer, membrane structure and use aerobic respiration to generate adenosi ...
l DNA studies conducted in 2004 on hundreds of samples from red deer and elk subspecies and other species of the ''Cervus'' deer family, strongly indicate that elk, or wapiti, should be a distinct species, namely ''Cervus canadensis''. DNA evidence validates that elk are more closely related to
Thorold's deer Thorold's deer (''Cervus albirostris'')Pitraa, Fickela, Meijaard, Groves (2004). ''Evolution and phylogeny of old world deer.'' Molecular Phylogenetics and Evolution 33: 880–895. is a threatened species of deer found in grassland, shrubland, an ...
and even
sika deer The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south to ...
than they are to the red deer. Elk and red deer produce fertile offspring in captivity, and the two species have freely inter-bred in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
's
Fiordland National Park Fiordland National Park occupies the southwest corner of the South Island of New Zealand. It is by far the largest of the 13 national parks in New Zealand, with an area of , and a major part of the Te Wahipounamu World Heritage Site. The park i ...
. The cross-bred animals have resulted in the disappearance of virtually all pure elk blood from the area. Key morphological differences that distinguish ''C. canadensis'' from ''C. elaphus'' are the former's wider rump patch and paler-hued antlers.


Subspecies

There are numerous
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of elk described, with six from North America and four from Asia, although some
taxonomists In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given ...
consider them different
ecotype In evolutionary ecology, an ecotype,Greek: ''οίκος'' = home and ''τύπος'' = type, coined by Göte Turesson in 1922 sometimes called ecospecies, describes a genetically distinct geographic variety, population, or race within a specie ...
s or races of the same species (adapted to local environments through minor changes in appearance and behavior). Populations vary in antler shape and size, body size, coloration and mating behavior. DNA investigations of the Eurasian subspecies revealed that
phenotypic In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological proper ...
variation in antlers, mane and rump patch development are based on "climatic-related lifestyle factors". Of the six subspecies of elk known to have inhabited North America in historical times, four remain, including the Roosevelt's (''C. canadensis roosevelti''),
tule ''Schoenoplectus acutus'' ( syn. ''Scirpus acutus, Schoenoplectus lacustris, Scirpus lacustris'' subsp. ''acutus''), called tule , common tule, hardstem tule, tule rush, hardstem bulrush, or viscid bulrush, is a giant species of sedge in the pl ...
(''C. canadensis nannodes''), Manitoban (''C. canadensis manitobensis'') and
Rocky Mountain elk The Rocky Mountain elk (''Cervus canadensis nelsoni'') is a subspecies of elk found in the Rocky Mountains and adjacent ranges of Western North America. Habitat The winter ranges are most common in open forests and floodplain marshes in the lo ...
(''C. canadensis nelsoni''). The
eastern elk The eastern elk (''Cervus canadensis canadensis'') is an extinct subspecies or distinct population of elk that inhabited the northern and eastern United States, and southern Canada. The last eastern elk was shot in Pennsylvania on September 1, 18 ...
(''C. canadensis canadensis'') and
Merriam's elk The Merriam's elk (''Cervus canadensis merriami'') is an extinct subspecies of elk once found in the arid lands of the southwestern United States (in Arizona, New Mexico and Texas), as well as in Mexico. Uncontrolled hunting and cattle grazing si ...
(''C. canadensis merriami'') subspecies have been extinct for at least a century. Four subspecies described in Asia include the Altai wapiti (''C. canadensis sibiricus'') and the Tianshan wapiti (''C. canadensis songaricus''). Two distinct subspecies found in
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
,
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
, the
Korean Peninsula Korea ( ko, 한국, or , ) is a peninsular region in East Asia. Since 1945, it has been divided at or near the 38th parallel, with North Korea (Democratic People's Republic of Korea) comprising its northern half and South Korea (Republic o ...
and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
are the
Manchurian wapiti The Manchurian wapiti (''Cervus canadensis xanthopygus'') is a subspecies of the wapiti native to East Asia. Description The Manchurian wapiti's coat is reddish brown during summer, and brownish gray in winter. It has dark hairs on the neck and ...
(''C. canadensis xanthopygus'') and the
Alashan wapiti The Alashan wapiti is a subspecies of ''Cervus canadensis'' (named " elk" or "wapiti" in North America), found in northern China and Mongolia. It is the smallest subspecies of elk, has the lightest color, and is the least studied, other than the ...
s (''C. canadensis alashanicus''). The
Manchuria Manchuria is an exonym (derived from the endo demonym " Manchu") for a historical and geographic region in Northeast Asia encompassing the entirety of present-day Northeast China (Inner Manchuria) and parts of the Russian Far East (Outer Manc ...
n wapiti is darker and more reddish in coloration than the other populations. The Alashan wapiti of north central China is the smallest of all subspecies, has the lightest coloration and is the least studied. Recent DNA studies suggest that there are no more than three or four subspecies of elk. All American forms, aside from possibly the tule and Roosevelt's elk, seem to belong to one subspecies (''Cervus canadensis canadensis''). Even the Siberian elk (''Cervus canadensis sibiricus'') are more or less identical to the American forms and therefore may belong to this subspecies, too. However, the Manchurian wapiti (''Cervus canadensis xanthopygus'') is clearly distinct from the Siberian forms, but not distinguishable from the Alashan wapiti. The Chinese forms (the
Sichuan deer The Sichuan deer (''Cervus canadensis macneilli''), also known as MacNeill's deer, is a subspecies of the Elk native to Western China. Description This large, highland deer is of very pale, finely spotted color with gray or brownish black. The ...
,
Kansu red deer The Kansu red deer (''Cervus canadensis kansuensis'') is a subspecies of wapiti found in the Gansu province of China. This subspecies forms, along with the closely related Sichuan deer, and Tibetan red deer The Tibetan red deer (''Cervus canad ...
, and
Tibetan red deer The Tibetan red deer (''Cervus canadensis wallichi'') also known as ''shou'', is a subspecies of elk/wapiti native to the southern Tibetan highlands and Bhutan. Once believed to be near-extinct, its population has increased to over 8,300, the maj ...
) also belong to the wapiti, and were not distinguishable from each other by mitochondrial DNA studies. These Chinese subspecies are sometimes treated as a distinct species, namely the
Central Asian red deer The Central Asian red deer (''Cervus hanglu''), also known as the Tarim red deer is a deer species native to Central Asia, where it used to be widely distributed, but is scattered today with small population units in several countries. It has bee ...
(''Cervus hanglu''), which also includes the
Kashmir stag The Kashmir stag (''Cervus hanglu hanglu''), also called hangul (), is a subspecies of Central Asian red deer endemic to Kashmir and surrounding areas. It is found in dense riverine forests in the high valleys and mountains of Jammu and Kashmi ...
.Mattioli (2011). ''Family Cervidae (Deer).'' (350-443). In: Wilson, D. E., Mittermeier, R. A., (Hrsg.). ''Handbook of the Mammals of the World. Volume 2: Hooved Mammals.'' Lynx Edicions, 2009. * North American group **
Roosevelt's elk The Roosevelt elk (''Cervus canadensis roosevelti)'', also known Common name, commonly as the Olympic elk and Roosevelt's wapiti, is the largest of the four surviving subspecies of elk (''Elk, Cervus canadensis'') in North America by body mass. ...
(''C. c. roosevelti'') **
Tule elk The tule elk (''Cervus canadensis nannodes'') is a subspecies of elk found only in California, ranging from the grasslands and marshlands of the Central Valley to the grassy hills on the coast. The subspecies name derives from the tule (), a s ...
(''C. c. nannodes'') **
Manitoban elk The Manitoban elk (''Cervus canadensis manitobensis'') is a subspecies of elk found in the Midwestern United States (specifically North Dakota) and southern regions of the Canadian Prairies (specifically Manitoba, Saskatchewan, and north-centr ...
(''C. c. manitobensis'') **
Rocky Mountain elk The Rocky Mountain elk (''Cervus canadensis nelsoni'') is a subspecies of elk found in the Rocky Mountains and adjacent ranges of Western North America. Habitat The winter ranges are most common in open forests and floodplain marshes in the lo ...
(''C. c. nelsoni'') **
Eastern elk The eastern elk (''Cervus canadensis canadensis'') is an extinct subspecies or distinct population of elk that inhabited the northern and eastern United States, and southern Canada. The last eastern elk was shot in Pennsylvania on September 1, 18 ...
(''C. c. canadensis''; extinct) **
Merriam's elk The Merriam's elk (''Cervus canadensis merriami'') is an extinct subspecies of elk once found in the arid lands of the southwestern United States (in Arizona, New Mexico and Texas), as well as in Mexico. Uncontrolled hunting and cattle grazing si ...
(''C. c. merriami''; extinct) * Eastern group ** Altai wapiti (''C. c. sibiricus'') **
Tian Shan wapiti The Tian Shan wapiti or Tian Shan maral (''Cervus canadensis songaricus''), is a subspecies of ''C. canadensis''. It is also called the Tian Shan elk in North American English. Description It is native to the Tian Shan Mountains in eastern Ky ...
(''C. c. songaricus'') **
Manchurian wapiti The Manchurian wapiti (''Cervus canadensis xanthopygus'') is a subspecies of the wapiti native to East Asia. Description The Manchurian wapiti's coat is reddish brown during summer, and brownish gray in winter. It has dark hairs on the neck and ...
(''C. c. xanthopygus'') **
Alashan wapiti The Alashan wapiti is a subspecies of ''Cervus canadensis'' (named " elk" or "wapiti" in North America), found in northern China and Mongolia. It is the smallest subspecies of elk, has the lightest color, and is the least studied, other than the ...
(''C. c. alashanicus'') **
Tibetan red deer The Tibetan red deer (''Cervus canadensis wallichi'') also known as ''shou'', is a subspecies of elk/wapiti native to the southern Tibetan highlands and Bhutan. Once believed to be near-extinct, its population has increased to over 8,300, the maj ...
(''C. c. wallichii'') File:Audubon-eastern-elk.jpg, Illustration of eastern elk File:The deer of all lands (1898) Altai wapiti.png, Illustration of Altai wapiti File:The deer of all lands (1898) Bedford's deer.png, Illustration of Manchurian wapiti File:The deer of all lands (1898) Hangul.png, Illustration of Kashmir stag


Characteristics

Elk have thick bodies with slender legs and short tails. They have a shoulder height of with a nose-to-tail length of . Males are larger and weigh while females weigh . The largest of the subspecies is the Roosevelt elk (''C. c. roosevelti''), found west of the
Cascade Range The Cascade Range or Cascades is a major mountain range of western North America, extending from southern British Columbia through Washington and Oregon to Northern California. It includes both non-volcanic mountains, such as the North Cascades, ...
in the
U.S. state In the United States, a state is a constituent political entity, of which there are 50. Bound together in a political union, each state holds governmental jurisdiction over a separate and defined geographic territory where it shares its sover ...
s of
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
,
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
and
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
, and in the
Canadian Canadians (french: Canadiens) are people identified with the country of Canada. This connection may be residential, legal, historical or cultural. For most Canadians, many (or all) of these connections exist and are collectively the source of ...
province of
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
. Roosevelt elk have been introduced into
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, where the largest males are estimated to weigh up to . More typically, male Roosevelt elk weigh around , while females weigh .. Male tule elk weigh while females weigh . The whole weights of adult male Manitoban elk range from . Females have a mean weight of . The elk is the second largest extant species of deer, after the
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
. Antlers are made of bone, which can grow at a rate of per day. While actively growing, a soft layer of highly vascularized skin known as
velvet Weave details visible on a purple-colored velvet fabric Velvet is a type of woven tufted fabric in which the cut threads are evenly distributed, with a short pile, giving it a distinctive soft feel. By extension, the word ''velvety'' means ...
covers and protects them. This is shed in the summer when the antlers have fully developed. Bull elk typically have around six tines on each antler. The Siberian and North American elk carry the largest antlers while the Altai wapiti has the smallest. Roosevelt bull antlers can weigh . The formation and retention of antlers are
testosterone Testosterone is the primary sex hormone and anabolic steroid in males. In humans, testosterone plays a key role in the development of Male reproductive system, male reproductive tissues such as testes and prostate, as well as promoting secondar ...
-driven. In late winter and early spring, the testosterone level drops, which causes the antlers to shed. During the fall, elk grow a thicker coat of hair, which helps to insulate them during the winter. Both male and female North American elk grow thin neck manes; females of other subspecies may not. By early summer, the heavy winter coat has been shed. Elk are known to rub against trees and other objects to help remove hair from their bodies. All elk have small and clearly defined rump patches with short tails. They have different coloration based on the seasons and types of habitats, with gray or lighter coloration prevalent in the winter and a more reddish, darker coat in the summer. Subspecies living in arid climates tend to have lighter colored coats than do those living in forests. Most have lighter yellow-brown to orange-brown coats in contrast to dark brown hair on the head, neck, and legs during the summer. Forest-adapted Manchurian and Alashan wapitis have red or reddish-brown coats with less contrast between the body coat and the rest of the body during the summer months. Calves are born spotted, as is common with many deer species, and lose them by the end of summer. Adult Manchurian wapiti may retain a few orange spots on the back of their summer coats until they are older. This characteristic has also been observed in the forest-adapted
European red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
.


Behavior and ecology

Elk are among the most
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
deer species. During the summer group size can reach 400 individuals. For most of the year, adult males and females are segregated into different herds. Female herds are larger while bulls form small groups and may even travel alone. Young bulls may associate with older bulls or female groups. Male and female herds come together during the mating season, which may begin in late August. Males try to intimidate rivals by vocalizing and displaying with their antlers. If neither bull backs down, they engage in antler wrestling, sometimes sustaining serious injuries. Bulls have a loud, high-pitched, whistle-like vocalization known as ''bugling'', which advertise the male's fitness over great distances. Unusual for a vocalization produced by a large animal, buglings can reach a frequency of 4000 Hz. This is achieved by blowing air from the
glottis The glottis is the opening between the vocal folds (the rima glottidis). The glottis is crucial in producing vowels and voiced consonants. Etymology From Ancient Greek ''γλωττίς'' (glōttís), derived from ''γλῶττα'' (glôtta), va ...
through the nasal cavities. Elk can produce deeper pitched (150 Hz) sounds using the
larynx The larynx (), commonly called the voice box, is an organ in the top of the neck involved in breathing, producing sound and protecting the trachea against food aspiration. The opening of larynx into pharynx known as the laryngeal inlet is about ...
. Cows produce an alarm bark to alert other members of the herd to danger, while calves will produce a high-pitched scream when attacked.


Reproduction and life cycle

Female elk have a short
estrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous p ...
cycle of only a day or two, and matings usually involve a dozen or more attempts. By the autumn of their second year, females can produce one and, very rarely, two offspring. Reproduction is most common when cows weigh at least . Dominant bulls follow groups of cows during the rut from August into early winter. A bull will defend his harem of 20 cows or more from competing bulls and predators. Bulls also dig holes in the ground called wallows, in which they urinate and roll their bodies. A male elk's urethra points upward so that urine is sprayed almost at a right angle to the penis. The urine soaks into their hair and gives them a distinct smell which attracts cows. A bull interacts with cows in his harem in two ways: herding and courtship. When a female wanders too far away from the harem's range, the male will rush ahead of her, block her path and aggressively rush her back to the harem. Herding behavior is accompanied by a stretched out and lowered neck and the antlers laid back. A bull may get violent and hit the cow with his antlers. During courtship, the bull is more peaceful and approaches her with his head and antlers raised. The male signals his intention to test the female for sexual receptivity by flicking his tongue. If not ready, a cow will lower her head and weave from side to side while opening and closing her mouth. The bull will stop in response in order not to scare her. Otherwise, the bull will copiously lick the female and then mount her. Younger, less dominant bulls, known as "spike bulls" because their antlers have not yet forked, will harass unguarded cows. These bulls are impatient and will not perform any courtship rituals and will continue to pursue a female even when she signals him to stop. As such, they are less reproductively successful, and a cow may stay close to the big bull to avoid harassment. Dominant bulls are intolerant of spike bulls and will chase them away from their harems. The
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
period is eight to nine months and the offspring weigh around . When the females are near to giving birth, they tend to isolate themselves from the main herd, and will remain isolated until the calf is large enough to escape predators. Calves are born spotted, as is common with many deer species, and they lose their spots by the end of summer. After two weeks, calves are able to join the herd, and are fully weaned at two months of age. Elk calves are as large as an adult
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
by the time they are six months old. Elk will leave their natal (birth) ranges before they are three years old. Males disperse more often than females, as adult cows are more tolerant of female offspring from previous years. Elk live 20 years or more in captivity but average 10 to 13 years in the wild. In some subspecies that suffer less predation, they may live an average of 15 years in the wild.


Migration

As is true for many species of deer, especially those in mountainous regions, elk migrate into areas of higher altitude in the spring, following the retreating snows, and the opposite direction in the fall. Hunting pressure impacts migration and movement. During the winter, they favor wooded areas for the greater availability of food to eat. Elk do not appear to benefit from thermal cover. The Greater Yellowstone Ecosystem elk herds comprise as many as 40,000 individuals. During the spring and fall, they take part in the longest elk migration in the continental U.S., traveling as much as between summer and winter ranges. The Teton herd consists of between 9,000 and 13,000 elk and they spend winters on the
National Elk Refuge The National Elk Refuge is a Wildlife Refuge located in Jackson Hole in the U.S. state of Wyoming. It was created in 1912 to protect habitat and provide sanctuary for one of the largest elk (also known as wapiti) herds. With a total of , the refu ...
, having migrated south from the southern portions of
Yellowstone National Park Yellowstone National Park is an American national park located in the western United States, largely in the northwest corner of Wyoming and extending into Montana and Idaho. It was established by the 42nd U.S. Congress with the Yellowston ...
and west from the
Shoshone The Shoshone or Shoshoni ( or ) are a Native American tribe with four large cultural/linguistic divisions: * Eastern Shoshone: Wyoming * Northern Shoshone: southern Idaho * Western Shoshone: Nevada, northern Utah * Goshute: western Utah, easter ...
and
Bridger–Teton National Forest Bridger–Teton National Forest is located in western Wyoming, United States. The forest consists of , making it the third largest National Forest outside Alaska. The forest stretches from Yellowstone National Park, along the eastern boundary ...
s.


Diet

Elk are ruminants and therefore have four-chambered stomachs. Unlike white-tailed deer and moose, which are chiefly browsers, elk are similar to
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
in that they are primarily grazers. But like other deer, they also
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
. Elk have a tendency to do most of their feeding in the mornings and evenings, seeking sheltered areas in between feedings to digest. Their diets vary somewhat depending on the season, with native grasses being a year-round supplement, tree bark being consumed in winter, and
forbs A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
and tree sprouts during the summer. Elk consume an average of of vegetation daily. Particularly fond of
aspen Aspen is a common name for certain tree species; some, but not all, are classified by botanists in the section ''Populus'', of the ''Populus'' genus. Species These species are called aspens: *'' Populus adenopoda'' – Chinese aspen (China ...
sprouts which rise in the spring, elk have had some impact on aspen groves which have been declining in some regions where elk exist. Range and wildlife managers conduct surveys of elk pellet groups to monitor populations and resource use.


Predators and defensive tactics

Predators of elk include
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
,
coyotes The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
,
brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing or painting, brown is usually made by combining the colors orange and black. In the RGB color model used ...
and
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have o ...
bears Bears are carnivoran mammals of the family Ursidae. They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats throughout the North ...
,
cougars The cougar (''Puma concolor'') is a large cat native to the Americas. Its range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mammal in the Western Hemisphere. I ...
, and
Siberian tigers The Siberian tiger or Amur tiger is a population of the tiger subspecies ''Panthera tigris tigris'' native to the Russian Far East, Northeast China and possibly North Korea. It once ranged throughout the Korean Peninsula, but currently inhabi ...
.
Coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
packs mostly prey on elk calves, though they can sometimes take a winter- or disease-weakened adult. In the
Greater Yellowstone Ecosystem The Greater Yellowstone Ecosystem (GYE) is one of the last remaining large, nearly intact ecosystems in the northern temperate zone of the Earth. It is located within the northern Rocky Mountains, in areas of northwestern Wyoming, southwestern Mo ...
, which includes Yellowstone National Park, bears are the most significant predators of calves while healthy bulls have never been recorded to be killed by bears and such encounters can be fatal for bears. The killing of cows in their prime is more likely to affect population growth than the killing of bulls or calves. Elk may avoid predation by switching from grazing to browsing. Grazing puts an elk in the compromising situation of being in an open area with its head down, leaving it unable to see what is going on in the surrounding area. Living in groups also lessens the risk of an individual falling to predation. Large bull elk are less vulnerable and can afford to wander alone, while cows stay in larger groups for protection for their calves. Bulls are more vulnerable to predation by wolves in late winter, after they have been weakened by months of chasing females and fighting. Males that have recently lost their antlers are more likely to be preyed upon.


Parasites and disease

At least 53 species of
protist A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the exc ...
and animal
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s have been identified in elk. Most of these parasites seldom lead to significant mortality among wild or captive elk. ''
Parelaphostrongylus tenuis ''Parelaphostrongylus tenuis'' (also known as meningeal worm or brainworm) is a neurotropic nematode parasite common to white-tailed deer, ''Odocoileus virginianus'', which causes damage to the central nervous system. Moose (''Alces alces'') ...
'' (brainworm or meningeal worm) is a parasitic
nematode The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-Parasitism, parasitic nematodes also known as eelworms. They are a diverse animal phylum inhab ...
known to affect the spinal cord and brain tissue of elk and other species, leading to death. The definitive host is the white-tailed deer, in which it normally has no ill effects. Snails and slugs, the intermediate hosts, can be inadvertently consumed by elk during grazing. The
liver fluke Liver fluke is a collective name of a polyphyletic group of parasitic trematodes under the phylum Platyhelminthes. They are principally parasites of the liver of various mammals, including humans. Capable of moving along the blood circulation, t ...
'' Fascioloides magna'' and the nematode ''
Dictyocaulus viviparus ''Dictyocaulus viviparus'' is a species of nematodes belonging to the family Dictyocaulidae. The species has cosmopolitan distribution In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all ...
'' are also commonly found parasites that can be fatal to elk. Since infection by either of these parasites can be lethal to some commercial livestock species, their presence in elk herds is of some concern.
Chronic wasting disease Chronic wasting disease (CWD), sometimes called zombie deer disease, is a transmissible spongiform encephalopathy (TSE) affecting deer. TSEs are a family of diseases thought to be caused by misfolded proteins called prions and include similar dis ...
, transmitted by a
misfolded protein Protein folding is the physical process by which a protein chain is translated to its native three-dimensional structure, typically a "folded" conformation by which the protein becomes biologically functional. Via an expeditious and reproduci ...
known as a
prion Prions are misfolded proteins that have the ability to transmit their misfolded shape onto normal variants of the same protein. They characterize several fatal and transmissible neurodegenerative diseases in humans and many other animals. It ...
, affects the brain tissue in elk, and has been detected throughout their range in North America. First documented in the late 1960s in mule deer, the disease has affected elk on game farms and in the wild in a number of regions. Elk that have contracted the disease begin to show weight loss, changes in behavior, increased watering needs, excessive salivation and urinating and difficulty swallowing, and at an advanced stage, the disease leads to death. No risks to humans have been documented, nor has the disease been demonstrated to pose a threat to domesticated cattle. In 2002,
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
banned the importation of elk antler velvet due to concerns about chronic wasting disease. The
Gram-negative Gram-negative bacteria are bacteria that do not retain the crystal violet stain used in the Gram staining method of bacterial differentiation. They are characterized by their cell envelopes, which are composed of a thin peptidoglycan cell wall ...
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ...
l disease
brucellosis Brucellosis is a highly contagious zoonosis caused by ingestion of unpasteurized milk or undercooked meat from infected animals, or close contact with their secretions. It is also known as undulant fever, Malta fever, and Mediterranean fever. The ...
occasionally affects elk in the Greater Yellowstone Ecosystem, the only place in the U.S. where the disease is still known to exist, though this can extend out to the
Bighorn Mountains The Bighorn Mountains ( cro, Basawaxaawúua, lit=our mountains or cro, Iisaxpúatahchee Isawaxaawúua, label=none, lit=bighorn sheep's mountains) are a mountain range in northern Wyoming and southern Montana in the United States, forming a nort ...
. In domesticated cattle, brucellosis causes infertility, abortions, and reduced milk production. It is transmitted to humans as undulant fever, producing
influenza Influenza, commonly known as "the flu", is an infectious disease caused by influenza viruses. Symptoms range from mild to severe and often include fever, runny nose, sore throat, muscle pain, headache, coughing, and fatigue. These symptoms ...
-like symptoms that may last for years. Though bison are more likely to transmit the disease to other animals, elk inadvertently transmitted brucellosis to horses in Wyoming and cattle in
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...
. Researchers are attempting to eradicate the disease through vaccinations and herd-management measures, which are expected to be successful. Nevertheless, research has been ongoing since 2002, and a successful vaccine has yet to be developed . A recent
necropsy An autopsy (post-mortem examination, obduction, necropsy, or autopsia cadaverum) is a surgical procedure that consists of a thorough examination of a corpse by dissection to determine the cause, mode, and manner of death or to evaluate any di ...
study of captive elk in Pennsylvania attributed the cause of death in 33 of 65 cases to either gastrointestinal parasites (21 cases, primarily ''
Eimeria ''Eimeria'' is a genus of apicomplexan parasites that includes various species capable of causing the disease coccidiosis in animals such as cattle, poultry and smaller ruminants including sheep and goats. ''Eimeria'' species are considered to be ...
'' sp. and ''
Ostertagia ''Ostertagia'' is a genus of nematodes belonging to the family Trichostrongylidae. The genus has cosmopolitan distribution. Species: *'' Ostertagia antipini'' *'' Ostertagia arctica'' *'' Ostertagia buriatica'' *'' Ostertagia dahurica'' *' ...
'' sp.) or bacterial infections (12 cases, mostly
pneumonia Pneumonia is an inflammatory condition of the lung primarily affecting the small air sacs known as alveoli. Symptoms typically include some combination of productive or dry cough, chest pain, fever, and difficulty breathing. The severity ...
). Elk hoof disease was first noticed in the state of Washington in the late 1990s in the
Cowlitz River The Cowlitz River is a river in the state of Washington in the United States, a tributary of the Columbia River. Its tributaries drain a large region including the slopes of Mount Rainier, Mount Adams, and Mount St. Helens. The Cowlitz has a ...
basin, with sporadic reports of deformed hooves. Since then, the disease has spread rapidly with increased sightings throughout southwest Washington and into Oregon. The disease is characterised by deformed, broken, or missing hooves and leads to severe lameness in elk. The primary cause is not known, but it is associated with treponeme bacteria, which are known to cause
digital dermatitis Digital dermatitis is a disease that causes lameness in cattle. It was first discovered in Italy in 1974 by Cheli and Mortellaro. This disease is caused by a mixture of different bacteria. Anaerobic bacteria, including spirochetes of the genus ...
in commercial livestock. The mode of transmission is also not known, but it appears to be highly contagious among elk. Studies are being undertaken by government departments to determine how to halt or eliminate the disease.


Distribution and status

The elk ranges from central Asia though to Siberia and east Asia and in North America. They can be found in open deciduous woodlands, boreal forests, upland moors, mountainous areas and grasslands. The
International Union for Conservation of Nature and Natural Resources The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) list the species as
least-concern species A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
. The habitat of Siberian elk in Asia is similar to that of the Rocky Mountain subspecies in North America. During the Late Pleistocene their range was much more extensive, being distributed across Eurasia, with remains being found as far west as France. These populations are most closely related to modern Asian populations of the elk. Their range collapsed at the start of the Holocene, possibly because they were specialized to cold periglacial tundra-steppe habitat. When this environment was replaced largely by closed forest the red deer might have outcompeted the elk. Relictual populations survived into the early Holocene (until around 3000 years ago) in southern Sweden and the Alps, where the environment remained favorable.


Introductions and reintroductions

As of 2014, population figures for all North American elk subspecies were around one million. Prior to the European colonization of North America, there were an estimated 10 million on the continent. There are many past and ongoing examples of reintroduction into areas of the US. Elk were reintroduced in
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and the ...
in 1918 after going extinct in 1875. The Rocky Mountain elk subspecies was reintroduced by hunter-conservation organizations into the
Appalachia Appalachia () is a cultural region in the Eastern United States that stretches from the Southern Tier of New York State to northern Alabama and Georgia. While the Appalachian Mountains stretch from Belle Isle in Newfoundland and Labrador, Ca ...
n region of the U.S. where the now extinct eastern elk once lived. Since the late 1990s, they were reintroduced and recolonized in the states of
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
,
Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...
,
North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia and So ...
,
Tennessee Tennessee ( , ), officially the State of Tennessee, is a landlocked state in the Southeastern region of the United States. Tennessee is the 36th-largest by area and the 15th-most populous of the 50 states. It is bordered by Kentucky to th ...
,
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
,
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
and
West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...
. In the state of Kentucky, the elk population in 2022 had increased to over 15,000 animals. In 2016, a male elk, likely from the
Smoky Mountains The Great Smoky Mountains (, ''Equa Dutsusdu Dodalv'') are a mountain range rising along the Tennessee–North Carolina border in the southeastern United States. They are a subrange of the Appalachian Mountains, and form part of the Blue Ridge ...
population, was sighted in
South Carolina )''Animis opibusque parati'' ( for, , Latin, Prepared in mind and resources, links=no) , anthem = " Carolina";" South Carolina On My Mind" , Former = Province of South Carolina , seat = Columbia , LargestCity = Charleston , LargestMetro = ...
for the first time in nearly 300 years. Since 2015, elk have also been reintroduced in a number of other states, including
Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...
,
Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ...
, and introduced to the islands of Etolin and
Afognak Afognak (Alutiiq language, Alutiiq: ''Agw’aneq''; russian: Афогнак) is an island in the Kodiak Archipelago north of Kodiak Island in the U.S. state of Alaska. It is long from east to west and wide from north to south and has a land ...
in Alaska. Reintroduction of the elk into Ontario began in the early 20th century and is ongoing with limited success. Elk and red deer were introduced to
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
in the early 20th century. There they are now considered an
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
, encroaching on Argentinian ecosystems where they compete for food with the indigenous
Chilean huemul The south Andean deer (''Hippocamelus bisulcus''), also known as the southern guemal, south Andean huemul, southern huemul, or Chilean ''huemul'' or '' güemul'' ( , ), is an endangered species of deer native to the mountains of Argentina and Chi ...
and other herbivores. This negative impact on native animal species has led the IUCN to identify the elk as one of the world's 100 worst invaders. The introduction of deer to
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
began in the middle of the 19th century, and current populations are primarily European red deer, with only 15 percent being elk. There is significant hybridization of elk with red deer. These deer have had an adverse impact on forest regeneration of some plant species, as they consume more palatable species, which are replaced with those that are less favored by the elk. The long-term impact will be an alteration of the types of plants and trees found, and in other animal and plant species dependent upon them. As in Chile and Argentina, the IUCN has declared that red deer and elk populations in New Zealand are an invasive species.


Estimated number of elk per U.S. state


Cultural references

Elk have played an important role in the cultural history of a number of peoples.
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
petroglyphs A petroglyph is an image created by removing part of a rock surface by incising, picking, carving, or abrading, as a form of rock art. Outside North America, scholars often use terms such as "carving", "engraving", or other descriptions ...
from Asia depict antler-less female elk, which have been interpreted as symbolizing life and sustenance. They were also frequently overlaid with boats and associated with rivers, suggesting they also represented paths to the underworld. Petroglyphs of elk were carved into cliffs by the
Ancestral Puebloans The Ancestral Puebloans, also known as the Anasazi, were an ancient Native American culture that spanned the present-day Four Corners region of the United States, comprising southeastern Utah, northeastern Arizona, northwestern New Mexico, a ...
of the southwestern U.S. hundreds of years ago. The elk was of particular importance to the
Lakota Lakota may refer to: *Lakota people, a confederation of seven related Native American tribes *Lakota language, the language of the Lakota peoples Place names In the United States: *Lakota, Iowa *Lakota, North Dakota, seat of Nelson County *Lakota ...
and played a spiritual role in their society. The male elk was admired for its ability to attract mates, and Lakota men will play a courting flute imitating a bugling elk to attract women. Men used elks' antlers as love charms and wore clothes decorated with elk images. The Rocky Mountain elk is the official state animal for
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
. An image of an elk and a moose appear on the state
seal Seal may refer to any of the following: Common uses * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impr ...
and
flag A flag is a piece of fabric (most often rectangular or quadrilateral) with a distinctive design and colours. It is used as a symbol, a signalling device, or for decoration. The term ''flag'' is also used to refer to the graphic design empl ...
of Michigan. The
Benevolent and Protective Order of Elks The Benevolent and Protective Order of Elks (BPOE; also often known as the Elks Lodge or simply The Elks) is an American fraternal order founded in 1868, originally as a social club in New York City. History The Elks began in 1868 as a soci ...
(B.P.O.E.) chose the elk as its namesake because a number of its attributes seemed appropriate for cultivation by members of the fraternity. Jewel encrusted, gold mounted elk teeth are prized possession of many members of the B.P.O.E..


Commercial uses

Although breakdown figures for each game species are not available in the 2006 National Survey from the U.S. Fish and Wildlife Service, hunting of wild elk is most likely the primary economic impact. While elk are not generally harvested for meat production on a large scale, some restaurants offer the meat as a specialty item and it is also available in some grocery stores. The meat has a taste somewhere between beef and
venison Venison originally meant the meat of a game animal but now refers primarily to the meat of antlered ungulates such as elk or deer (or antelope in South Africa). Venison can be used to refer to any part of the animal, so long as it is edible, in ...
and is higher in
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respo ...
and lower in
fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers spec ...
and
cholesterol Cholesterol is any of a class of certain organic molecules called lipids. It is a sterol (or modified steroid), a type of lipid. Cholesterol is biosynthesized by all animal cells and is an essential structural component of animal cell mem ...
than beef, pork, and chicken. Elk meat is a good source of
iron Iron () is a chemical element with symbol Fe (from la, ferrum) and atomic number 26. It is a metal that belongs to the first transition series and group 8 of the periodic table. It is, by mass, the most common element on Earth, right in f ...
,
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
and
zinc Zinc is a chemical element with the symbol Zn and atomic number 30. Zinc is a slightly brittle metal at room temperature and has a shiny-greyish appearance when oxidation is removed. It is the first element in group 12 (IIB) of the periodi ...
. A male elk can produce of antler velvet annually and on ranches in the United States, Canada and New Zealand, it is collected and sold to markets in East Asia, where it is used in medicine. Some cultures consider velvet to be an
aphrodisiac An aphrodisiac is a substance that increases sexual desire, sexual attraction, sexual pleasure, or sexual behavior. Substances range from a variety of plants, spices, foods, and synthetic chemicals. Natural aphrodisiacs like cannabis or cocain ...
. However, consuming velvet from elk in North America may be risky since velvet from animals infected with chronic wasting disease may contain
prions Prions are misfolded proteins that have the ability to transmit their misfolded shape onto normal variants of the same protein. They characterize several fatal and transmissible neurodegenerative diseases in humans and many other animals. It i ...
that could result in a human getting
variant Creutzfeldt–Jakob disease Variant Creutzfeldt–Jakob disease (vCJD), commonly referred to as "mad cow disease" or "human mad cow disease" to distinguish it from its BSE counterpart, is a fatal type of brain disease within the transmissible spongiform encephalopathy fa ...
. Antlers are also used in artwork, furniture and other novelty items. All Asian subspecies, along with other deer, have been raised for their antlers in central and eastern Asia by
Han Chinese The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctive va ...
,
Turkic peoples The Turkic peoples are a collection of diverse ethnic groups of West, Central, East, and North Asia as well as parts of Europe, who speak Turkic languages.. "Turkic peoples, any of various peoples whose members speak languages belonging t ...
,
Tungusic peoples Tungusic peoples are an ethno-linguistic group formed by the speakers of Tungusic languages (or Manchu–Tungus languages). They are native to Siberia and Northeast Asia. The Tungusic phylum is divided into two main branches, northern (Evenic or ...
,
Mongolians The Mongols ( mn, Монголчууд, , , ; ; russian: Монголы) are an East Asian ethnic group native to Mongolia, Inner Mongolia in China and the Buryatia Republic of the Russian Federation. The Mongols are the principal member of ...
, and
Koreans Koreans ( South Korean: , , North Korean: , ; see names of Korea) are an East Asian ethnic group native to the Korean Peninsula. Koreans mainly live in the two Korean nation states: North Korea and South Korea (collectively and simply refe ...
. Elk farms are relatively common in North America and New Zealand. Native Americans have used elk hides for
tepee A tipi , often called a lodge in English, is a conical tent, historically made of animal hides or pelts, and in more recent generations of canvas, stretched on a framework of wooden poles. The word is Siouan, and in use in Dakhótiyapi, Lakȟó ...
covering, clothing and footwear. Since 1967, the
Boy Scouts of America The Boy Scouts of America (BSA, colloquially the Boy Scouts) is one of the largest scouting organizations and one of the largest youth organizations in the United States, with about 1.2 million youth participants. The BSA was founded i ...
have assisted employees at the
National Elk Refuge The National Elk Refuge is a Wildlife Refuge located in Jackson Hole in the U.S. state of Wyoming. It was created in 1912 to protect habitat and provide sanctuary for one of the largest elk (also known as wapiti) herds. With a total of , the refu ...
in Wyoming by collecting the antlers which are shed each winter. They are then auctioned, with 80% of the proceeds returned to the refuge. In 2010, of antlers were auctioned, bringing in over $46,000.


References


External links


Arizona Elk
Arizona Game and Fish Department The Arizona Game and Fish Department is a state agency of Arizona, headquartered in Phoenix. The agency is tasked with conserving, enhancing, and restoring Arizona's diverse wildlife resources and habitats through protection and management progra ...

Rocky Mountain Elk FoundationSmithsonian Institution - North American Mammals: ''Cervus'' (''elaphus'') ''canadensis''
{{Taxonbar, from=Q180404 Cervus Elk and red deer Fauna of Siberia Holarctic fauna Mammals of the United States Fauna of Central Asia Mammals of Canada Fauna of East Asia Extant Pleistocene first appearances Herbivorous mammals Mammals described in 1777 Taxa named by Johann Christian Polycarp Erxleben